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In the actions editor conditions (IF, ELSE IF, ELSE) and actions (like Create issue, Add comment, Execute transition, Auto replyAutoreply, etc...) can be added. Conditions and action actions can be added under each other and are executed in that order they appear in the list on the left side. 

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Actions added below a condition are executed if the given IF or ELSE condition is evaluated to true. There are some special actions, like Set field value that can be added under other actions like Create issue or Execute transition actions. In such cases, these child actions will be executed together with the parent action in a single step. For example, when a new issue is created, fields can be initialized as well, like a single transaction, avoiding to generate a few notification emails about issue updated in that case these actions were executed separatedlyseparately, each in a single step.

To add an action under a condition or another action, hover the mouse and use the plus ( + ) sign on the right to open the pop-up menu:

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The following is a very simple mail handler:
It checks if issue is not found (by issue keys found in the email's subject) and creates a new Bug issue in SP project. The issue's summary will be the email's subject and the description will contain the email's body.
Else, if the issue was found, then a comment will be added to the existing issue and the comment's content will be the email's body.

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